* {
    margin: 0;
    padding: 0;
    box-sizing: border-box;
}

:root {
    --color-background-primary: #F7F9F7;
    --color-text-primary: #2E3D35;
    --color-accent-forest: #225740;
    --color-background-sage: #E8EDE9;
    --color-border-subtle: #D4DDD6;
    --color-hover-tint: #1A4532;
    --font-serif: 'Playfair Display', Georgia, serif;
    --font-sans: 'Inter', -apple-system, BlinkMacSystemFont, 'Segoe UI', sans-serif;
}

body {
    font-family: var(--font-sans);
    font-size: 16px;
    line-height: 1.7;
    color: var(--color-text-primary);
    background-color: var(--color-background-primary);
    font-weight: 400;
}

.encyclopedia-masthead {
    background-color: #fff;
    border-bottom: 2px solid var(--color-border-subtle);
    padding: 2rem 0;
    margin-bottom: 3rem;
}

.container-primary {
    max-width: 1400px;
    margin: 0 auto;
    padding: 0 2rem;
}

.branding-zone {
    display: flex;
    align-items: baseline;
    gap: 1.5rem;
}

.publication-title {
    font-family: var(--font-serif);
    font-size: 2.5rem;
    font-weight: 700;
    color: var(--color-accent-forest);
    letter-spacing: -0.02em;
}

.taxonomic-subtitle {
    font-size: 0.875rem;
    font-weight: 300;
    color: var(--color-text-primary);
    opacity: 0.7;
    text-transform: uppercase;
    letter-spacing: 0.05em;
}

.dual-column-layout {
    display: grid;
    grid-template-columns: 1fr 380px;
    gap: 3rem;
    margin-bottom: 4rem;
}

.content-primary-stream {
    min-width: 0;
}

.navigation-compact {
    margin-bottom: 2rem;
}

.menu-expandable {
    background-color: #fff;
    border: 1px solid var(--color-border-subtle);
    border-radius: 4px;
    padding: 0;
}

.menu-expandable summary {
    padding: 0.875rem 1.25rem;
    cursor: pointer;
    font-weight: 500;
    color: var(--color-accent-forest);
    user-select: none;
    list-style: none;
    transition: background-color 0.2s;
}

.menu-expandable summary::-webkit-details-marker {
    display: none;
}

.menu-expandable summary::before {
    content: '▸ ';
    display: inline-block;
    margin-right: 0.5rem;
    transition: transform 0.2s;
}

.menu-expandable[open] summary::before {
    transform: rotate(90deg);
}

.menu-expandable summary:hover {
    background-color: var(--color-background-sage);
}

.menu-expandable ul {
    list-style: none;
    padding: 0.5rem 1.25rem 1.25rem 1.25rem;
    border-top: 1px solid var(--color-border-subtle);
}

.menu-expandable ul li {
    padding: 0.5rem 0;
}

.menu-expandable ul li a {
    color: var(--color-text-primary);
    text-decoration: none;
    transition: color 0.2s;
}

.menu-expandable ul li a:hover {
    color: var(--color-accent-forest);
}

.encyclopedia-entry {
    background-color: #fff;
    padding: 2.5rem;
    border-radius: 6px;
    box-shadow: 0 1px 3px rgba(0, 0, 0, 0.05);
    margin-bottom: 2rem;
}

.encyclopedia-entry h1,
.encyclopedia-entry h2,
.encyclopedia-entry h3,
.encyclopedia-entry h4,
.encyclopedia-entry h5,
.encyclopedia-entry h6 {
    font-family: var(--font-serif);
    color: var(--color-accent-forest);
    line-height: 1.3;
    margin-bottom: 1rem;
}

.encyclopedia-entry h1 {
    font-size: 2.25rem;
    font-weight: 700;
    margin-bottom: 1.5rem;
    border-bottom: 2px solid var(--color-border-subtle);
    padding-bottom: 1rem;
}

.encyclopedia-entry h2 {
    font-size: 1.75rem;
    font-weight: 600;
    margin-top: 2.5rem;
}

.encyclopedia-entry h3 {
    font-size: 1.35rem;
    font-weight: 600;
    margin-top: 2rem;
}

.encyclopedia-entry p {
    margin-bottom: 1.25rem;
}

.encyclopedia-entry ul,
.encyclopedia-entry ol {
    margin: 1.25rem 0;
    padding-left: 2rem;
}

.encyclopedia-entry a {
    color: var(--color-accent-forest);
    text-decoration: none;
    border-bottom: 1px solid var(--color-border-subtle);
    transition: all 0.2s;
}

.encyclopedia-entry a:hover {
    color: var(--color-hover-tint);
    border-bottom-color: var(--color-hover-tint);
}

.phenotype-gallery {
    background-color: #fff;
    padding: 2.5rem;
    border-radius: 6px;
    box-shadow: 0 1px 3px rgba(0, 0, 0, 0.05);
}

.section-nomenclature {
    font-family: var(--font-serif);
    font-size: 1.75rem;
    font-weight: 600;
    color: var(--color-accent-forest);
    margin-bottom: 1.5rem;
    padding-bottom: 0.75rem;
    border-bottom: 2px solid var(--color-border-subtle);
}

.specimen-grid {
    display: grid;
    grid-template-columns: repeat(auto-fit, minmax(180px, 1fr));
    gap: 1.5rem;
}

.specimen-card {
    background-color: var(--color-background-sage);
    border-radius: 4px;
    overflow: hidden;
    transition: transform 0.2s, box-shadow 0.2s;
}

.specimen-card:hover {
    transform: translateY(-2px);
    box-shadow: 0 4px 12px rgba(34, 87, 64, 0.15);
}

.specimen-card img {
    width: 100%;
    height: 200px;
    object-fit: cover;
    display: block;
}

.specimen-card figcaption {
    padding: 1rem;
    font-size: 0.875rem;
    font-weight: 500;
    color: var(--color-text-primary);
    text-align: center;
}

.sidebar-supplemental {
    display: flex;
    flex-direction: column;
    gap: 2rem;
}

.lexicon-module,
.nutrient-chart-module,
.reference-module {
    background-color: var(--color-background-sage);
    padding: 1.75rem;
    border-radius: 6px;
    border: 1px solid var(--color-border-subtle);
}

.module-header {
    font-family: var(--font-serif);
    font-size: 1.25rem;
    font-weight: 600;
    color: var(--color-accent-forest);
    margin-bottom: 1.25rem;
    padding-bottom: 0.75rem;
    border-bottom: 1px solid var(--color-border-subtle);
}

.terminology-list {
    display: flex;
    flex-direction: column;
    gap: 1rem;
}

.lexicon-term {
    font-weight: 600;
    color: var(--color-accent-forest);
    font-size: 0.95rem;
    margin-bottom: 0.25rem;
}

.term-definition {
    font-size: 0.875rem;
    line-height: 1.6;
    color: var(--color-text-primary);
}

.growth-phase-container {
    display: flex;
    flex-direction: column;
    gap: 1.5rem;
}

.phase-block {
    padding: 1rem;
    background-color: #fff;
    border-radius: 4px;
}

.phase-label {
    font-family: var(--font-sans);
    font-size: 0.95rem;
    font-weight: 600;
    color: var(--color-accent-forest);
    margin-bottom: 0.75rem;
}

.npk-indicator {
    display: flex;
    flex-direction: column;
    gap: 0.5rem;
    margin-bottom: 0.75rem;
}

.nutrient-bar {
    display: grid;
    grid-template-columns: 30px 1fr;
    align-items: center;
    gap: 0.5rem;
}

.element-symbol {
    font-weight: 600;
    font-size: 0.85rem;
    color: var(--color-text-primary);
}

.bar-fill {
    height: 8px;
    background-color: var(--color-accent-forest);
    border-radius: 4px;
    transition: width 0.3s;
}

.bar-fill.high-level {
    width: 100%;
}

.bar-fill.medium-level {
    width: 60%;
}

.bar-fill.low-level {
    width: 25%;
}

.phase-description {
    font-size: 0.8rem;
    line-height: 1.5;
    color: var(--color-text-primary);
    opacity: 0.85;
}

.taxonomy-table {
    width: 100%;
    font-size: 0.875rem;
}

.taxonomy-table tr {
    display: grid;
    grid-template-columns: 90px 1fr;
    padding: 0.5rem 0;
    border-bottom: 1px solid var(--color-border-subtle);
}

.taxonomy-table tr:last-child {
    border-bottom: none;
}

.taxonomy-table th {
    text-align: left;
    font-weight: 600;
    color: var(--color-accent-forest);
}

.taxonomy-table td {
    color: var(--color-text-primary);
    font-style: italic;
}

.encyclopedia-footer {
    background-color: #fff;
    border-top: 2px solid var(--color-border-subtle);
    padding: 2rem 0;
    margin-top: 4rem;
}

.footer-content {
    display: flex;
    flex-direction: column;
    gap: 0.5rem;
    align-items: center;
    text-align: center;
}

.contact-information,
.copyright-notice {
    font-size: 0.875rem;
    color: var(--color-text-primary);
    opacity: 0.8;
}

.contact-information a {
    color: var(--color-accent-forest);
    text-decoration: none;
    transition: color 0.2s;
}

.contact-information a:hover {
    color: var(--color-hover-tint);
}

@media (max-width: 1024px) {
    .dual-column-layout {
        grid-template-columns: 1fr;
    }

    .sidebar-supplemental {
        order: -1;
    }
}

@media (max-width: 768px) {
    .container-primary {
        padding: 0 1rem;
    }

    .publication-title {
        font-size: 1.75rem;
    }

    .branding-zone {
        flex-direction: column;
        gap: 0.5rem;
    }

    .encyclopedia-entry,
    .phenotype-gallery {
        padding: 1.5rem;
    }

    .specimen-grid {
        grid-template-columns: repeat(auto-fit, minmax(140px, 1fr));
        gap: 1rem;
    }

    .specimen-card img {
        height: 150px;
    }
}